- UU – Utrecht University (the Coordinator)
- EMBL – European Molecular Biology Laboratory
- Diamond – Diamond Light Source
- CIRMMP – Consorzio Interuniversitario Risonanze Magnetiche di Metallo Proteine
- BMRZ – Goethe University Frankfurt
- NKI – The Netherlands Cancer Institute (the Deputy Coordinator)
- CEITEC – Masaryk University / CEITEC
- CSIC – Spanish National Research Council
- FVB-FMP – Leibniz-Institut für Molekulare Pharmakologie
- SOLEIL – Synchrotron SOLEIL
- MAXIV – Lund University / MAXIV
- NeCEN – Leiden University / NeCEN
- LUMC – Leiden University Medical Center
- ESRF – European Synchrotron Radiation Facility
- RALF-NMR – Centre National de la Recherche Scientifique / RALF-NMR
- Instruct – ESFRI Instruct
- AU – Aarhus University
- ESS – European Spallation Source
- ELTE – Eötvös Loránd University
- ITQB – Instituto de Tecnologia Química e Biológica
- OULU – University of Oulu
- UPAT – University of Patras
- Weizmann – Weizmann Institute of Science
- Instruct (ESFRI) – Pan-European Research Infrastructure from the ESFRI Roadmap, providing expertise and access to high quality instruments for structural cell biology researchers. Instruct is also a partner in iNEXT.
- ESS (ESFRI) – The large European Spallation Source (ESS) research infrastructure (RI) from the ESFRI Roadmap will be a world first in the field of neutron production for analytical purposes. The ESS is also a partner in iNEXT.
- EuroBioImaging (ESFRI) – The ESFRI research infrastructure Euro-BioImaging will provide open user access to a complete range of state-of-the-art imaging technologies in biological, molecular and medical imaging for life scientists in Europe and beyond.
- EU-OPENSCREEN (ESFRI) – EU-OPENSCREEN is building a sustainable European infrastructure for Chemical Biology, supporting life science research and its translation to medicine, agriculture, bioindustries and society, as part of the ESFRI Roadmap.